39. Rhodostemonodaphne recurva

Rhodostemonodaphne recurva van der Werff

Novon 10: 291. 2000. Type. Brazil. Amazonas: Mun. Manaus, Reserva Florestal Ducke, 5 Aug 1994 (stam. fl), Vicentini et al. 653 (holotype, INPA n.v.; isotypes, MO, SP n.v.).

Description

Trees: branches basitonic, in axils of cataphylls; twigs angular, soon becoming terete, ca. 3 mm diam.; epidermis brownish to black, barely visible due to indument cover; terminal bud plump, ca. 5 X 3 mm; cataphylls caducous; indument sericeous, caducous by next flush, the hairs dense, up to 0.25 mm long, straight, appressed ascending to erect, golden yellow. Leaves: petioles slender, 1.5–3 cm X 1.5–2.5 mm, adaxially flattened; blades chartaceous, flat, widely elliptic to oblong (obovate), 6–14 X 3–5.5 cm; base cuneate to acute, minutely decurrent, 65–80°; apex rounded, 80–100°, mucronate up to 1 cm long; margin recurved at base or raised; primary vein above flat, below prominent; secondary veins 3–4 pairs, equidistant, eucamptodromous, above flat, below slightly raised, diverging at ca. 30°, gradually arching, chordal angle ca. 20°, the angle increasing towards apex; tertiary veins above and below slightly raised, random-reticulate to scalariform; higher order veins above and below flat; surface above shiny brown, below dull brown (grayish on young leaves); indument above absent, below minutely sericeous, the hairs sparse, up to 0.4 mm long, straight, appressed, ascending, yellowish to translucent, persistent. Staminate inflorescences: along whole length of flush, erect, peduncles (4–)5–7 cm long, the hypopodia 2.5–3.5 cm X ca. 1 mm, branch orders 4, the second-order branches 3–5, dispersed, lowest branch up to 2.5 cm long, all axes color indument as on twigs; bracts and bracteoles caducous (not seen). Staminate flowers: pedicels ca. 2.5 X 0.6 mm, gradually enlarging to form an obconical receptacle ca. 1.5 X 2 mm; tepals membranaceous, ovate, ca. 1.7 X 2 mm (inner whorl slightly larger), at anthesis spreading, yellowish, adaxially puberulous; stamens of whorls I and II, the anthers sessile, globose, ca. 0.8 X 1.3 mm (whorl II slightly smaller), glabrous, the locelli (1,) 2 (3, 4), apical, in a shallow arch, introrse; whorl III columnar, ca. 1 X 0.7 mm, glabrous, the anthers columnar, the locelli 4, the upper pair latrorse, the lower pair extrorse, the glands 2, minute ca. 2 mm broad, black; whorl IV absent; all stamens reddish; pistillode minute, elongate, ca. 2 mm long. Pistillate plant not seen.

Field notes

Trees 17–25 m tall and 30–35 cm diam.; trunk straight; outer bark smooth with inconspicuous elliptic to roundish lenticels arranged in vertical lines and alternating with longitudinal fissures; <1 mm thick; inner bark two layered, the external beige, homogeneous, the internal mixed dark and light beige; wood whitish yellow.  Inflorescence axis green with reddish branchlets and pedicels; tepals whitish green adaxially, yellow green abaxially.

Distribution (Figure 23)

Rhodostemonodaphne recurva is only known from the surroundings of Manaus.  It grows on terra firme in yellow clay soils.

Additional specimens examined

Brazil. Amazonas: Distrito agropecuário de SUFRAMA, Rodovia BR-174, Km 72, Fazenda Dimona, 6 Ago 1982 (stam. fl), Mackenzie (INPA/WWF 2108/35) (MO); ibid. Km 64, Fazenda Esteio, 2 Oct 1980 (stam. fl), J. L. dos Santos (INPA/WWF 1202/5767) (MO); ibid. s.d. (stam. fl) s.coll. (INPA/WWF 1202/3094) (MO).

Discussion

When I first saw the specimens that now represent R. recurva, I determined them as an undescribed species of Endlicheria due to their two-locellate anthers.  The plants however share a number of characters with the species in the R. grandis-complex, such as the sericeous indument, and the flowers with spreading tepals and broad sessile anthers.

Contact | Updated 29.07.2005 | ©2005 Santiago Madriñán