19. Rhodostemonodaphne curicuriariensis

Rhodostemonodaphne curicuriariensis Madriñán, sp. nov.

Type. Brazil. Amazonas: Rio Negro, above mouth of Curicuriari river, 24 Oct 1932 (pist. fl), Ducke 25674 (holotype, U).

Frutex insignis, petiolis crassis in sicco nigrescentibus, foliis glabris in sicco flavoviridibus, bullatis, venatione supra elevata a congeneris diversa.

Named after the Curicuriari mountain range and river where the two known specimens were collected.

Description

Scandent? shrubs: branching unknown; twigs angular and remaining so for at least two flushes, ca. 3 mm diam.; epidermis yellowish to brownish; terminal bud slender, 4 X 2 mm; cataphylls caducous; indument tomentose, caducous by next flush, the hairs sparse, up to 0.4 mm long, straight, appressed, ascending, yellowish. Leaves: petioles robust, ca. 1.5 cm X 3.5 mm, conspicuously blackened, terete; blades chartaceous, flat to undulate to bullate, narrowly elliptic, 17–31 X 7–9 cm; base acute, ca. 70°; apex acute to obtuse, 70–90°, acuminate for up to 2 cm; margin plane; primary vein above and below raised; secondary veins 7–9 pairs, with distinct intersecondaries, equidistant, brochidodromous, above slightly raised, below raised, diverging at ca. 60°, evenly arching, chordal angle ca. 35°, the angle uniform along blade length; tertiary veins above slightly impressed, below slightly raised, random-reticulate; higher order veins above inconspicuous, below slightly raised; surface above shiny light brownish-green, below greenish-brown; indument above absent, below puberulous, soon glabrescent, the hairs isolated, up to 0.4 mm long, straight, appressed, ascending, yellowish. Staminate plant unknown. Pistillate inflorescences: acrotonic?, pendulous?, peduncles ca. 15 cm long, the hypopodia ca. 5 cm X 1 mm, branch orders 3, the second-order branches 11, dispersed, lowest branch up to ca. 1.7 cm long, color and indument of all axes as on twigs; bracts caducous, up to 3 mm long, adaxially hairy; bracteoles persistent, up to 0.8 mm long, adaxially hairy. Pistillate flowers: pedicels ca. 2.8 X 0.6 mm, the diameter even throughout; receptacle obconical, ca. 1.2 X 2.4 mm; tepals membranaceous, elliptic to oblong, ca. 2 X 1.6 mm, at anthesis spreading, reddish to yellowish-brown, adaxially glabrous; staminodes of whorls I and II, the anthers sessile chubby, trapezoid, ca. 1 X 1.4 mm (whorl II slightly smaller), glabrous, the locelli 4, apical, in a shallow arch, introrse, the glands absent; whorl III columnar, ca. 1.2 X 1.2 mm, glabrous, the anthers trapezoid, the locelli 4, the upper pair latrorse, the lower pair extrorse, the glands (if present) each pair fused to each other and the base of the stamen which is thickened; whorl IV absent; all staminodes reddish-brown to black; pistil ca. 2.5 X 1.6 mm; ovary globose, ca. 1.6 mm long, glabrous. Fruits unknown.

Field notes

Shrubs? Tepals green.

Distribution (Figure 11)

Known from only two specimens from the Curicuriari mountains near the mouth of the river with the same name, a tributary of the Rio Negro.  Growing in non-inundated forest at ca. 100 m elev.  The two flowering specimens were collected in October (ca. 50 years apart), at the beginning of the dry season.

Additional specimens examined

Brazil. Amazonas: Rio Negro, above mouth of Curicuriari river, Curucuriari mountains, 24 Oct 1976 (pist. fl), Nascimento 735 (NY).

Discussion

Rhodostemonodaphne curicuriariensis has leaves with distinctly blackened petioles, similar to those found in Endlicheria rubriflora Mez and some other species of Lauraceae.  The leaves are conspicuously shiny and glabrous above and the yellow veins are raised from an impressed groove.  The two specimens cited, although undoubtedly conspecific, have very different leaf blades. In the type these are flat to slightly undulate, while in Nascimento 735 they are distinctly bullate.  This situation is not uncommon in the genus (see “Morphology”).

Contact | Updated 29.07.2005 | ©2005 Santiago Madriñán